Subfamily NACOBBODERINAE

                                Rev 4/91

Classification:

      Tylenchida
       Tylenchina
        Tylenchoidea
         Heteroderidae
          Nacobboderinae (Golden & Jensen, 1974)
    Synonyms:
      Meloinematinae (Meloineminae) Husain, 1976

Morphology and Anatomy:

      Cuticle not abnormally thick, annulated in all stages of males and 
      females.  Cephalic framework strongly sclerotized.  Stylet robust. 
      Female: Pre-adult female vermiform, free in the substrate, with two 
      opposed immature genital branches and a short tail.  Mature female 
      sedentary, spheroid/pear-shaped, with very long neck and swollen body 
      exposed.  Labial area set-off, with irregular annulation (plates) and a 
      reduced non-offset labial disc.  Vulva subterminal, well removed from
      anus; no finger-like print pattern around vulva.  Phasmids conspicuous.  
      Remnants of tail present.  No cyst stage.  Eggs not retained in female
      body, but deposited in a gelatinous matrix.  Male: Labial area high, 
      annulated and set-off.  Lateral field with four or five lines.  Tail 
      conoid-rounded, without caudal alae (exception: Brusadera).  Males 
      develop within second-stage cuticle.  Juveniles: Cephalic
      sclerotization and stylet robust.  Lateral field with four lines.  
      Biology: Sedentary, causing galls on roots.
      [Ref: Luc, Maggenti & Fortuner, Rev. Nematol. 11(2):159-176 (1988).]

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